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By interweaving Hegelian dialectic and the middle voice, this book develops a holistic account of life, nature, and the ethical orientation of human beings with respect to them, without falling into the trap of either subjecting human rights to totality or relegating non-human beings and their habitats to instrumentalism. Both ontological problems that have arisen in the conceptualization of non-human life as well as ethical problems that arise within human relations to non-human living beings and their ecosystems are addressed. Presenting a systematic derivation of categories, life is shown to be irreducible to mechanico-chemical processes and that, taken together with the sphere of rights and obligations unique to human beings, the specific character of the category of life implies an ecological ethic whose object of concern includes species as well as habitats. The ontological character of life is in turn clarified in terms of the middle voice, providing a philosophically grounded human-nature holism that preserves the specific differences important for both inter-human and ecological ethics on the one hand and for both philosophy and the empirical sciences on the other.